Reference health
The bHLH network underlying plant shade‐avoidance
86/86 checkable references clean · checked 2026-07-22
Every reference with a DOI in the deposited reference list resolved to a known work in Crossref or DataCite at the dated check, and none carried a retraction, withdrawal, or removal notice.
2 without a DOI — not checked. A reference deposited without a DOI is never matched by title or guessed at; it stays outside the checked set, and this line discloses that.
The 86 checked references that resolve
resolves10.1105/tpc.112.105163
A Triple Helix-Loop-Helix/Basic Helix-Loop-Helix Cascade Controls Cell Elongation Downstream of Multiple Hormonal and Environmental Signaling Pathways in<i>Arabidopsis</i>
resolves10.1038/ncb2546
Brassinosteroid, gibberellin and phytochrome impinge on a common transcription module in Arabidopsis
resolves10.1105/tpc.151140
Update on the Basic Helix-Loop-Helix Transcription Factor Gene Family in <i>Arabidopsis thaliana</i>
resolves10.1104/pp.110.153593
Genome-Wide Classification and Evolutionary Analysis of the bHLH Family of Transcription Factors in Arabidopsis, Poplar, Rice, Moss, and Algae
resolves10.1111/j.1399-3054.2012.01600.x
The non‐<i>DNA</i>‐binding b<i>HLH</i> transcription factor <i>PRE3/bHLH135/ATBS1/TMO7</i> is involved in the regulation of light signaling pathway in <i>Arabidopsis</i>
resolves10.1111/tpj.12264
The b<scp>HLH</scp> proteins <scp>BEE</scp> and <scp>BIM</scp> positively modulate the shade avoidance syndrome in <scp>A</scp>rabidopsis seedlings
resolves10.1111/jipb.12822
Photoexcited phytochrome B interacts with brassinazole resistant 1 to repress brassinosteroid signaling in <i>Arabidopsis</i>
resolves10.1016/j.cub.2004.12.026
The Degradation of HFR1, a Putative bHLH Class Transcription Factor Involved in Light Signaling, Is Regulated by Phosphorylation and Requires COP1
resolves10.1038/emboj.2009.4
Interaction of COP1 and UVR8 regulates UV‐B‐induced photomorphogenesis and stress acclimation in Arabidopsis
resolves10.1093/genetics/162.3.1445
Three Redundant Brassinosteroid Early Response Genes Encode Putative bHLH Transcription Factors Required for Normal Growth
resolves10.1105/tpc.16.00790
Molecular Profiles of Contrasting Shade Response Strategies in Wild Plants: Differential Control of Immunity and Shoot Elongation
resolves10.1016/j.cub.2016.10.001
Shade Promotes Phototropism through Phytochrome B-Controlled Auxin Production
resolves10.1093/mp/sss011
Interactions between HLH and bHLH Factors Modulate Light-Regulated Plant Development
resolves10.1073/pnas.1403052111
UV-B detected by the UVR8 photoreceptor antagonizes auxin signaling and plant shade avoidance
resolves10.1126/science.1107580
BZR1 Is a Transcriptional Repressor with Dual Roles in Brassinosteroid Homeostasis and Growth Responses
resolves10.1093/molbev/msg088
The Basic Helix-Loop-Helix Transcription Factor Family in Plants: A Genome-Wide Study of Protein Structure and Functional Diversity
resolves10.1073/pnas.1320355111
Light intensity modulates the regulatory network of the shade avoidance response in
<i>Arabidopsis</i>
resolves10.1007/s10059-013-2159-2
A Competitive Peptide Inhibitor KIDARI Negatively Regulates HFR1 by Forming Nonfunctional Heterodimers in Arabidopsis Photomorphogenesis
resolves10.1038/emboj.2009.306
Inhibition of the shade avoidance response by formation of non‐DNA binding bHLH heterodimers
resolves10.1111/j.1365-313X.2012.05033.x
Phytochrome interacting factors 4 and 5 control seedling growth in changing light conditions by directly controlling auxin signaling
resolves10.1007/s11103-006-0010-2
KIDARI, Encoding a Non-DNA Binding bHLH Protein, Represses Light Signal Transduction in Arabidopsis thaliana
resolves10.1105/tpc.112.105023
A Triantagonistic Basic Helix-Loop-Helix System Regulates Cell Elongation in <i>Arabidopsis</i>
resolves10.4161/psb.23448
ATBS1 INTERACTING FACTORs negatively regulate<i>Arabidopsis</i>cell elongation in the triantagonistic bHLH system
resolves10.1111/j.1365-313X.2011.04598.x
Cryptochrome 1 and phytochrome B control shade‐avoidance responses in Arabidopsis via partially independent hormonal cascades
resolves10.1111/j.1365-313X.2011.04597.x
Blue‐light‐mediated shade avoidance requires combined auxin and brassinosteroid action in Arabidopsis seedlings
resolves10.1111/tpj.12423
Darkness and <i>gulliver2</i>/<i>phy<scp>B</scp></i> mutation decrease the abundance of phosphorylated <scp>BZR</scp>1 to activate brassinosteroid signaling in <scp>A</scp>rabidopsis
resolves10.1105/tpc.16.00463
Neighbor Detection Induces Organ-Specific Transcriptomes, Revealing Patterns Underlying Hypocotyl-Specific Growth
resolves10.1093/pcp/pcj026
Overexpression of PRE1 and its Homologous Genes Activates Gibberellin-dependent Responses in Arabidopsis thaliana
resolves10.1093/mp/ssu068
PIL1 Participates in a Negative Feedback Loop that Regulates Its Own Gene Expression in Response to Shade
resolves10.1016/j.devcel.2017.12.028
UVR8 Interacts with BES1 and BIM1 to Regulate Transcription and Photomorphogenesis in Arabidopsis
resolves10.1073/pnas.1700850114
Noncanonical role of
<i>Arabidopsis</i>
COP1/SPA complex in repressing BIN2-mediated PIF3 phosphorylation and degradation in darkness
resolves10.1126/science.1163927
Photoexcited CRY2 Interacts with CIB1 to Regulate Transcription and Floral Initiation in
<i>Arabidopsis</i>
resolves10.1111/j.1365-313X.2007.03341.x
Phytochrome‐mediated inhibition of shade avoidance involves degradation of growth‐promoting bHLH transcription factors
resolves10.1242/dev.152892
Regulation of intercellular TARGET OF MONOPTEROS 7 protein transport in the
<i>Arabidopsis</i>
root
resolves10.1105/tpc.113.121657
COP1 and phyB Physically Interact with PIL1 to Regulate Its Stability and Photomorphogenic Development in<i>Arabidopsis</i>
resolves10.1073/pnas.1511437113
Cryptochrome 1 interacts with PIF4 to regulate high temperature-mediated hypocotyl elongation in response to blue light
resolves10.1111/nph.16194
Photoexcited CRY1 and phyB interact directly with ARF6 and ARF8 to regulate their DNA‐binding activity and auxin‐induced hypocotyl elongation in <i>Arabidopsis</i>
resolves10.1105/tpc.109.065946
The<i>Arabidopsis</i>Floral Homeotic Proteins APETALA3 and PISTILLATA Negatively Regulate the<i>BANQUO</i>Genes Implicated in Light Signaling
resolves10.15252/embj.201899552
PIF4‐induced BR synthesis is critical to diurnal and thermomorphogenic growth
resolves10.1073/pnas.0407107101
The phytochrome-interacting transcription factor, PIF3, acts early, selectively, and positively in light-induced chloroplast development
resolves10.1146/annurev.genet.39.073003.112240
Concerted and Birth-and-Death Evolution of Multigene Families
resolves10.1016/S0092-8674(00)81636-0
PIF3, a Phytochrome-Interacting Factor Necessary for Normal Photoinduced Signal Transduction, Is a Novel Basic Helix-Loop-Helix Protein
resolves10.1038/ncb2545
Interaction between BZR1 and PIF4 integrates brassinosteroid and environmental responses
resolves10.7554/eLife.03031
Cell elongation is regulated through a central circuit of interacting transcription factors in the Arabidopsis hypocotyl
resolves10.1111/nph.13965
Convergence of <scp>CONSTITUTIVE PHOTOMORPHOGENESIS</scp> 1 and <scp>PHYTOCHROME INTERACTING FACTOR</scp> signalling during shade avoidance
resolves10.1111/j.1365-313X.2012.05114.x
Phytochrome B inhibits binding of phytochrome‐interacting factors to their target promoters
resolves10.1016/j.cell.2015.12.018
Cryptochromes Interact Directly with PIFs to Control Plant Growth in Limiting Blue Light
resolves10.1093/mp/ssu087
Combinatorial Complexity in a Transcriptionally Centered Signaling Hub in Arabidopsis
resolves10.1093/jxb/ert389
Shade avoidance: phytochrome signalling and other aboveground neighbour detection cues
resolves10.1104/pp.108.133496
Auxin and Ethylene Regulate Elongation Responses to Neighbor Proximity Signals Independent of Gibberellin and DELLA Proteins in Arabidopsis
resolves10.1016/j.pbi.2018.04.018
Photoreceptor-mediated regulation of the COP1/SPA E3 ubiquitin ligase
resolves10.1104/pp.105.076331
Identification of Primary Target Genes of Phytochrome Signaling. Early Transcriptional Control during Shade Avoidance Responses in Arabidopsis
resolves10.1038/sj.emboj.7601890
Interaction of shade avoidance and auxin responses: a role for two novel atypical bHLH proteins
resolves10.1016/j.biotechadv.2011.08.014
Plant adaptation to dynamically changing environment: The shade avoidance response
resolves10.1038/nature08836
MONOPTEROS controls embryonic root initiation by regulating a mobile transcription factor
resolves10.1101/gad.364005
A dynamic balance between gene activation and repression regulates the shade avoidance response in
<i>Arabidopsis</i>
resolves10.1038/s41467-019-12369-1
UVR8 disrupts stabilisation of PIF5 by COP1 to inhibit plant stem elongation in sunlight
resolves10.1111/j.1365-313X.2012.04992.x
Phytochrome‐imposed oscillations in PIF3 protein abundance regulate hypocotyl growth under diurnal light/dark conditions in Arabidopsis
resolves10.1093/jxb/ert465
PIF1 promotes phytochrome-regulated growth under photoperiodic conditions in Arabidopsis together with PIF3, PIF4, and PIF5
resolves10.1111/tpj.14556
Degradation of the transcription factors PIF4 and PIF5 under UV‐B promotes UVR8‐mediated inhibition of hypocotyl growth in Arabidopsis
resolves10.1105/tpc.109.072504
Regulation of<i>Arabidopsis</i>Brassinosteroid Signaling by Atypical Basic Helix-Loop-Helix Proteins
resolves10.1093/jxb/ery209
CRY1 interacts directly with HBI1 to regulate its transcriptional activity and photomorphogenesis in Arabidopsis
resolves10.1105/tpc.17.00994
Photoexcited CRYPTOCHROME1 Interacts with Dephosphorylated BES1 to Regulate Brassinosteroid Signaling and Photomorphogenesis in Arabidopsis
resolves10.1111/nph.15866
BES1‐regulated BEE1 controls photoperiodic flowering downstream of blue light signaling pathway in <i>Arabidopsis</i>
resolves10.1016/j.cub.2016.10.031
Integration of Phytochrome and Cryptochrome Signals Determines Plant Growth during Competition for Light
resolves10.1093/pcp/pcy212
phyB Interacts with BES1 to Regulate Brassinosteroid Signaling in Arabidopsis
resolves10.1242/dev.146936
Reciprocal proteasome-mediated degradation of PIFs and HFR1 underlies photomorphogenic development in <i>Arabidopsis</i>
resolves10.1016/j.cell.2004.11.044
A New Class of Transcription Factors Mediates Brassinosteroid-Regulated Gene Expression in Arabidopsis
resolves10.1111/j.1365-313X.2010.04449.x
A brassinosteroid transcriptional network revealed by genome‐wide identification of BESI target genes in <i>Arabidopsis thaliana</i>
resolves10.1105/tpc.109.070441
Antagonistic HLH/bHLH Transcription Factors Mediate Brassinosteroid Regulation of Cell Elongation and Plant Development in Rice and<i>Arabidopsis</i>
resolves10.3389/fpls.2017.01813
Involvement of PACLOBUTRAZOL RESISTANCE6/KIDARI, an Atypical bHLH Transcription Factor, in Auxin Responses in Arabidopsis
resolves10.1073/pnas.1400203111
Helix–loop–helix/basic helix–loop–helix transcription factor network represses cell elongation in
<i>Arabidopsis</i>
through an apparent incoherent feed-forward loop
resolves10.1104/pp.113.227231
Both PHYTOCHROME RAPIDLY REGULATED1 (PAR1) and PAR2 Promote Seedling Photomorphogenesis in Multiple Light Signaling Pathways
The 2 references without a DOI — listed, not checked
no DOI — not checkedShade avoidance
no DOI — not checkedHFR1 encodes an atypical bHLH protein that acts in phytochrome a signal transduction
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