Reference health

The 3D architecture of the ctenophore aboral organ and the evolution of complex integrative centers in animals

https://doi.org/10.1126/sciadv.aea8399
CiteStamped reference-health badge
55/55 checkable references clean · checked 2026-08-20

Every reference with a DOI in the deposited reference list resolved to a known work in Crossref or DataCite at the dated check, and none carried a retraction, withdrawal, or removal notice.

6 without a DOI — not checked. A reference deposited without a DOI is never matched by title or guessed at; it stays outside the checked set, and this line discloses that.

The 55 checked references that resolve
resolves10.1111/brv.12239
The origin of the animals and a ‘Savannah’ hypothesis for early bilaterian evolution
resolves10.1016/j.conb.2021.11.002
Evolution of synapses and neurotransmitter systems: The divide-and-conquer model for early neural cell-type evolution
resolves10.1038/nrn.2015.15
From nerve net to nerve ring, nerve cord and brain — evolution of the nervous system
resolves10.3389/fevo.2020.00082
Of Circuits and Brains: The Origin and Diversification of Neural Architectures
resolves10.1038/s41467-021-22074-7
Evidence for sponges as sister to all other animals from partitioned phylogenomics with mixture models and recoding
resolves10.1016/j.cub.2017.02.031
A Large and Consistent Phylogenomic Dataset Supports Sponges as the Sister Group to All Other Animals
resolves10.1073/pnas.1518127112
Genomic data do not support comb jellies as the sister group to all other animals
resolves10.1371/journal.pbio.3002632
The promise and pitfalls of synteny in phylogenomics
resolves10.1093/molbev/msaf321
Sponges, ctenophores, and the statistical significance of syntenies
resolves10.1038/nature06614
Broad phylogenomic sampling improves resolution of the animal tree of life
resolves10.1038/nature13400
The ctenophore genome and the evolutionary origins of neural systems
resolves10.1126/science.1242592
The Genome of the Ctenophore <i>Mnemiopsis leidyi</i> and Its Implications for Cell Type Evolution
resolves10.1038/s41586-023-05936-6
Ancient gene linkages support ctenophores as sister to other animals
resolves10.1016/j.tree.2015.03.003
The hidden biology of sponges and ctenophores
resolves10.1073/pnas.2122052119
Ctenophores are direct developers that reproduce continuously beginning very early after hatching
resolves10.1111/ivb.12042
Cilia and the life of ctenophores
resolves10.5962/bhl.title.39844
Die ctenophoren des golfes von Neapel, und der angrenzenden meeres-abschnitte. Eine monographie von Carl Chun.
resolves10.1126/science.ade5645
Syncytial nerve net in a ctenophore adds insights on the evolution of nervous systems
resolves10.1007/978-3-642-51593-4_9
Ctenophora
resolves10.1242/jcs.s3-105.71.311
Presumed photoreceptive cilia in a ctenophore
resolves10.1111/j.1525-142X.2005.05051.x
Larval and adult brains <sup>1</sup>
resolves10.1186/1741-7007-12-7
Larval body patterning and apical organs are conserved in animal evolution
resolves10.1093/molbev/msad285
Gene Regulatory Network that Shaped the Evolution of Larval Apical Organ in Cnidaria
resolves10.1016/j.ydbio.2024.02.010
Graded FGF activity patterns distinct cell types within the apical sensory organ of the sea anemone Nematostella vectensis
resolves10.1139/z02-130
Metamorphosis in the Cnidaria
resolves10.3389/fendo.2020.00063
Expression Analysis of Cnidarian-Specific Neuropeptides in a Sea Anemone Unveils an Apical-Organ-Associated Nerve Net That Disintegrates at Metamorphosis
resolves10.1002/cbin.12023
The ultrastructure of the apical organ of the Müller's larva of the tiger flatworm <i>Prostheceraeus crozieri</i>
resolves10.1126/sciadv.adg6034
Single-cell atlases of two lophotrochozoan larvae highlight their complex evolutionary histories
resolves10.1073/pnas.1220285110
Conserved MIP receptor–ligand pair regulates <i>Platynereis</i> larval settlement
resolves10.1371/journal.pbio.1001488
The Bilaterian Head Patterning Gene six3/6 Controls Aboral Domain Development in a Cnidarian
resolves10.1242/dev.200833
Molecular and cellular architecture of the larval sensory organ in the cnidarian <i>Nematostella vectensis</i>
resolves10.1186/2041-9139-1-10
Genomic insights into Wnt signaling in an early diverging metazoan, the ctenophore Mnemiopsis leidyi
resolves10.1186/2041-9139-1-9
The homeodomain complement of the ctenophore Mnemiopsis leidyi suggests that Ctenophora and Porifera diverged prior to the ParaHoxozoa
resolves10.1002/jmor.10019
Novel bridge of axon‐like processes of epithelial cells in the aboral sense organ of ctenophores
resolves10.1093/icb/5.3.357
RELATIONS BETWEEN NERVES AND CILIA IN CTENOPHORES
resolves10.1093/database/baaa029
The Mnemiopsis Genome Project Portal: integrating new gene expression resources and improving data visualization
resolves10.1186/2041-9139-5-4
A cleavage clock regulates features of lineage-specific differentiation in the development of a basal branching metazoan, the ctenophore Mnemiopsis leidyi
resolves10.7554/eLife.108420
Neural connectome of the ctenophore statocyst
resolves10.1086/BBLv229n2p173
Functional Consequences of the Asymmetric Architecture of the Ctenophore Statocyst
resolves10.1016/j.cub.2021.09.005
Neuropeptide repertoire and 3D anatomy of the ctenophore nervous system
resolves10.1242/jeb.114.1.443
Unilateral Ciliary Reversal and Motor Responses During Prey Capture by the Ctenophore <i>Pleurobrachia</i>
resolves10.1126/sciadv.abo2416
Homologous gene regulatory networks control development of apical organs and brains in Bilateria
resolves10.1111/dgd.12812
Gene activation of metazoan Fox transcription factors at the onset of metamorphosis in the marine demosponge <i>Amphimedon queenslandica</i>
resolves10.1016/j.cub.2023.07.022
An RFX transcription factor regulates ciliogenesis in the closest living relatives of animals
resolves10.1007/978-1-0716-3642-8_4
Stable Laboratory Culture System for the Ctenophore Mnemiopsis leidyi
resolves10.1038/nmeth.2019
Fiji: an open-source platform for biological-image analysis
resolves10.1371/journal.pone.0038011
TrakEM2 Software for Neural Circuit Reconstruction
resolves10.1186/1471-2105-11-274
A high-level 3D visualization API for Java and ImageJ
resolves10.1186/s12983-021-00414-z
Improved histological fixation of gelatinous marine invertebrates
resolves10.1242/dev.165753
Third-generation <i>in situ</i> hybridization chain reaction: multiplexed, quantitative, sensitive, versatile, robust
resolves10.3389/fphys.2022.882413
Optimization of Whole Mount RNA Multiplexed in situ Hybridization Chain Reaction With Immunohistochemistry, Clearing and Imaging to Visualize Octopus Embryonic Neurogenesis
resolves10.1093/molbev/msaa159
Broccoli: Combining Phylogenetic and Network Analyses for Orthology Assignment
resolves10.1093/nar/gkh340
MUSCLE: multiple sequence alignment with high accuracy and high throughput
resolves10.1093/bioinformatics/btp348
trimAl: a tool for automated alignment trimming in large-scale phylogenetic analyses
resolves10.1093/molbev/msaa015
IQ-TREE 2: New Models and Efficient Methods for Phylogenetic Inference in the Genomic Era
The 6 references without a DOI — listed, not checked
no DOI — not checkedR. Hertwig Über Den Bau Der Ctenophoren - Studien Zur Blättertheorie (G. Fischer 1880).
no DOI — not checkedS. L. Tamm “Ctenophora” in Electrical Conduction and Behavior in “Simple” Invertebrates (Shelton G. A. B. 1982) pp. 266–358.
no DOI — not checkedL. Leclère, M. Bause, C. Sinigaglia, J. Steger, F. Rentzsch, Development of the aboral domain in Nematostella requires β-catenin and the opposing activities of six3/6 and frizzled5/8. Development 143, 1766–1777 (2016).
no DOI — not checkedM.-L. Hernandez-Nicaise “Système nerveux et intégration chez les Cténaires. Etude ultrastructurale et comportementale ” thesis Univ. Claude Bernard Lyon France (1974).
no DOI — not checkedM. Z. Aronova, Electron microscopy observation on the aboral organ of Ctenophora. I. The gravity receptor. Z. Mikrosk. Anat. Forsch. 88, 401–412 (1974).
no DOI — not checkedA. Jan M. Candelas J. J. Soto-Angel “Métodos de cultivo de zooplancton gelatinoso” in Zooplancton Gelatinoso (El Colegio de la Frontera Sur ed. 1 2026).
What this badge says. CiteStamped means the CHECKABLE references of this work were clean at the dated check: each resolved to a known work in a public registry, and none carried a retraction notice at that time. It says nothing about the quality, findings, or importance of the work itself, and nothing about references deposited without a DOI.

checked 2026-08-20 — re-checked daily as this page is visited; titles and statuses come from Crossref and DataCite and are not part of the signed record

Embed this badge

Both snippets point at the live badge image and link back to this page. The badge re-renders from the daily check, so an embed never goes stale by more than a day of visits.

<a href="https://citestamp.com/citestamped/10.1126/sciadv.aea8399"><img src="https://citestamp.com/citestamped/10.1126/sciadv.aea8399/badge.svg" alt="CiteStamped reference-health badge" width="460" height="64"></a>
[![CiteStamped reference-health badge](https://citestamp.com/citestamped/10.1126/sciadv.aea8399/badge.svg)](https://citestamp.com/citestamped/10.1126/sciadv.aea8399)