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The Tempo and Mode of Three‐Dimensional Morphological Evolution in Male Reproductive Structures

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What this work says

Outgoing claims: the papers this work cites, builds on, or argues with.

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Efficient Registration of 3D SPHARM Surfaces

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CORRELATED EVOLUTION OF MALE AND FEMALE MORPHOLOGIES IN WATER STRIDERS

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MATE RECOGNITION AND MECHANICAL ISOLATION IN ENALLAGMA DAMSELFLIES (ODONATA: COENAGRIONIDAE)

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LIFETIME REPRODUCTIVE SUCCESS AND THE OPPORTUNITY FOR SELECTION IN A NONTERRITORIAL DAMSELFLY (ODONATA: COENAGRIONIDAE)

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PHYLOGENETIC ANALYSES OF THE CORRELATED EVOLUTION OF CONTINUOUS CHARACTERS: A SIMULATION STUDY

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LABORATORY EXPERIMENTS ON SPECIATION: WHAT HAVE WE LEARNED IN 40 YEARS?

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INTRASEXUAL COMPETITION ALONE FAVORS A SEXUALLY DIMORPHIC ORNAMENT IN THE RUBYSPOT DAMSELFLYHETAERINA AMERICANA

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SEXUAL SELECTION, SPACE, AND SPECIATION

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BIOCHEMICAL EVOLUTION ASSOCIATED WITH ANTIPREDATOR ADAPTATION IN DAMSELFLIES

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ADAPTIVE GAMETE-RECOGNITION DIVERGENCE IN A HYBRIDIZING MYTILUS POPULATION

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Incoming claims: where this work appears in other papers’ reference lists and arguments.

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inferred← citeshttps://doi.org/10.1016/j.cag.2009.03.002openalex-snapshot

Fourier method for large-scale surface modeling and registration

inferred← citeshttps://doi.org/10.1016/j.cub.2009.06.056openalex-snapshot

Evolutionary Response to Sexual Selection in Male Genital Morphology

inferred← citeshttps://doi.org/10.1016/j.forsciint.2017.06.016openalex-snapshot

Voyaging into the third dimension: A perspective on virtual methods and their application to studies of juvenile sex estimation and the ontogeny of sexual dimorphism

inferred← citeshttps://doi.org/10.1016/j.powtec.2022.117461openalex-snapshot

Estimation of three-dimensional particle size and shape characteristics using a modified 2D–3D conversion method employing spherical harmonic-based principal component analysis

inferred← citeshttps://doi.org/10.1016/j.powtec.2022.117462openalex-snapshot

Applicable scope of estimation of three-dimensional particle size distribution by the Goldsmith-Cruz-Orive and improved methods

inferred← citeshttps://doi.org/10.1016/j.powtec.2022.118143openalex-snapshot

Reproducibility of the repose angle, porosity, and coordination number of particles generated by spherical harmonic-based principal component analysis using discrete element simulation

inferred← citeshttps://doi.org/10.1016/j.powtec.2024.119488openalex-snapshot

Generation of 3D realistic geological particles using conditional generative adversarial network aided spherical harmonic analysis

inferred← citeshttps://doi.org/10.1016/j.powtec.2025.121109openalex-snapshot

3D particle shape estimation without pre-training or prior models

inferred← citeshttps://doi.org/10.1016/j.tree.2021.03.003openalex-snapshot

When Ecology Fails: How Reproductive Interactions Promote Species Coexistence

inferred← citeshttps://doi.org/10.1016/j.ympev.2015.08.005openalex-snapshot

Asymmetry in genitalia does not increase the rate of their evolution

inferred← citeshttps://doi.org/10.1016/j.ympev.2017.06.008openalex-snapshot

Convergent and unidirectional evolution of extremely long aedeagi in the largest feather mite genus, Proctophyllodes (Acari: Proctophyllodidae): Evidence from comparative molecular and morphological phylogenetics

inferred← citeshttps://doi.org/10.1017/pab.2024.31openalex-snapshot

Punctuated equilibrium: state of the evidence

inferred← citeshttps://doi.org/10.1017/pab.2024.35openalex-snapshot

Morphological evolution in a time of phenomics

inferred← citeshttps://doi.org/10.1038/nrg2829openalex-snapshot

Evolution and development of shape: integrating quantitative approaches

inferred← citeshttps://doi.org/10.1038/s41437-024-00719-9openalex-snapshot

Reproductive character displacement: insights from genital morphometrics in damselfly hybrid zones

inferred← citeshttps://doi.org/10.1038/s41437-026-00837-6openalex-snapshot

Alternative reproductive strategies explain asymmetries in reproductive isolation and reinforcement in two Ischnura damselfly species

inferred← citeshttps://doi.org/10.1061/(asce)mt.1943-5533.0003851openalex-snapshot

Aggregate Geometry Generation Method Using a Structured Light 3D Scanner, Spherical Harmonics–Based Geometry Reconstruction, and Placing Algorithms for Mesoscale Modeling of Concrete

inferred← citeshttps://doi.org/10.1080/08912963.2024.2350556openalex-snapshot

A new species of Eocene soldier beetle, a Malthodes (Coleoptera: Cantharidae: Malthininae) in Baltic amber

inferred← citeshttps://doi.org/10.1080/13887890.2011.651983openalex-snapshot

Morphometric and molecular studies on the populations of the damselfliesChalcolestes viridisandC. parvidens(Odonata, Lestidae)

inferred← citeshttps://doi.org/10.1080/19336934.2018.1429859openalex-snapshot

Combining morphology and molecular data to improveDrosophila paulistorum(Diptera, Drosophilidae) taxonomic status

inferred← citeshttps://doi.org/10.1086/593137openalex-snapshot

The Ecological Dynamics of Clade Diversification and Community Assembly

inferred← citeshttps://doi.org/10.1086/657060openalex-snapshot

Adaptation andThe Origin of Species

inferred← citeshttps://doi.org/10.1086/673206openalex-snapshot

Ecology and Sexual Selection: Evolution of Wing Pigmentation in Calopterygid Damselflies in Relation to Latitude, Sexual Dimorphism, and Speciation

inferred← citeshttps://doi.org/10.1086/691101openalex-snapshot

The Ecological Dynamics of Natural Selection: Traits and the Coevolution of Community Structure

inferred← citeshttps://doi.org/10.1086/697201openalex-snapshot

Frequency Dependence and Ecological Drift Shape Coexistence of Species with Similar Niches

inferred← citeshttps://doi.org/10.1086/713664openalex-snapshot

Accelerated Brain Shape Evolution Is Associated with Rapid Diversification in an Avian Radiation

inferred← citeshttps://doi.org/10.1093/beheco/arag028openalex-snapshot

Intraspecific variation in animal mating signals: a test of Mayr's conjecture

inferred← citeshttps://doi.org/10.1093/bioinformatics/btu315openalex-snapshot

Ceiba: scalable visualization of phylogenies and 2D/3D image collections

inferred← citeshttps://doi.org/10.1093/bioinformatics/btu335openalex-snapshot

RiboMaker: computational design of conformation-based riboregulation

inferred← citeshttps://doi.org/10.1093/biolinnean/blad008openalex-snapshot

Ornamentation diversified faster than eco-morphology across Nearctic dragonflies

inferred← citeshttps://doi.org/10.1093/biolinnean/blae123openalex-snapshot

Macroevolutionary and biomechanical implications of rostral flexion in bat skulls: a major early driver of cranial evolution in bats

inferred← citeshttps://doi.org/10.1093/biolinnean/blx073openalex-snapshot

Genetic and phenotypic differentiation suggests incipient speciation within Drosophila arizonae (Diptera: Drosophilidae)

inferred← citeshttps://doi.org/10.1093/biolinnean/bly108openalex-snapshot

Rapid divergence of courtship song in the face of neutral genetic homogeneity in the cactophilic fly Drosophila buzzatii

inferred← citeshttps://doi.org/10.1093/evolut/qpaf253openalex-snapshot

Coincident transitions across elevation and origins of functional innovations drove the phenotypic and ecological diversity of lungless salamanders

inferred← citeshttps://doi.org/10.1093/isd/ixaf022openalex-snapshot

The evolution of 2 male terminalia periphallic structures in Drosophila and related genera (Diptera: Drosophilidae)

inferred← citeshttps://doi.org/10.1093/isd/ixz019openalex-snapshot

Toward Phylomics in Entomology: Current Systematic and Evolutionary Morphology

inferred← citeshttps://doi.org/10.1093/oso/9780192864161.003.0008openalex-snapshot

Interspecific hybridization in insects in times of climate change

inferred← citeshttps://doi.org/10.1093/sysbio/syac058openalex-snapshot

Functional Trade-Offs Asymmetrically Promote Phenotypic Evolution

inferred← citeshttps://doi.org/10.1093/sysbio/syp106openalex-snapshot

Testing and Quantifying Phylogenetic Signals and Homoplasy in Morphometric Data

inferred← citeshttps://doi.org/10.1093/sysbio/sys083openalex-snapshot

Comparing Evolutionary Rates for Different Phenotypic Traits on a Phylogeny Using Likelihood

inferred← citeshttps://doi.org/10.1093/sysbio/syt105openalex-snapshot

Quantifying and Comparing Phylogenetic Evolutionary Rates for Shape and Other High-Dimensional Phenotypic Data

inferred← citeshttps://doi.org/10.1093/sysbio/syu030openalex-snapshot

A Generalized K Statistic for Estimating Phylogenetic Signal from Shape and Other High-Dimensional Multivariate Data

inferred← citeshttps://doi.org/10.1098/rsos.221603openalex-snapshot

Interspecific hybridization explains rapid gorget colour divergence inHeliodoxahummingbirds (Aves: Trochilidae)

inferred← citeshttps://doi.org/10.1098/rspb.2009.0543openalex-snapshot

Are rates of species diversification correlated with rates of morphological evolution?

inferred← citeshttps://doi.org/10.1098/rspb.2014.1636openalex-snapshot

Sex differences in developmental plasticity and canalization shape population divergence in mate preferences

inferred← citeshttps://doi.org/10.1098/rspb.2014.1737openalex-snapshot

Interspecific aggression, not interspecific mating, drives character displacement in the wing coloration of male rubyspot damselflies ( Hetaerina )

inferred← citeshttps://doi.org/10.1098/rspb.2018.2014openalex-snapshot

Metabolic physiology explains macroevolutionary trends in the melanic colour system across amniotes

inferred← citeshttps://doi.org/10.1098/rspb.2023.2883openalex-snapshot

Rapid divergent evolution of internal female genitalia and the coevolution of male genital morphology revealed by micro-computed tomography

inferred← citeshttps://doi.org/10.1098/rstb.2017.0418openalex-snapshot

Sexual conflict in its ecological setting

inferred← citeshttps://doi.org/10.1101/2021.03.08.434376openalex-snapshot

Sexual conflict promotes species coexistence through negative frequency dependence

inferred← citeshttps://doi.org/10.1101/2022.05.16.492149openalex-snapshot

Choice of 3D morphometric method leads to diverging interpretations of form–function relationships in the carnivoran calcaneus

inferred← citeshttps://doi.org/10.1101/2023.10.26.564279openalex-snapshot

Reproductive character displacement explains strengthening of mechanical barriers in damselflies

inferred← citeshttps://doi.org/10.1109/bibm55620.2022.9995036openalex-snapshot

Using Optimal Transport to Improve Spherical Harmonic Quantification of Complex Biological Shapes

inferred← citeshttps://doi.org/10.1111/2041-210x.12085openalex-snapshot

Testing the link between phenotypic evolution and speciation: an integrated palaeontological and phylogenetic analysis

inferred← citeshttps://doi.org/10.1111/2041-210x.12954openalex-snapshot

A new method for testing evolutionary rate variation and shifts in phenotypic evolution

inferred← citeshttps://doi.org/10.1111/aen.12053openalex-snapshot

Sexual selection and genital evolution

inferred← citeshttps://doi.org/10.1111/bij.12347openalex-snapshot

Rapid evolution of prezygotic barriers in non-territorial damselflies

inferred← citeshttps://doi.org/10.1111/cla.12599openalex-snapshot

Incomplete barriers to heterospecific mating among Somatochlora species (Odonata: Corduliidae) as revealed in multi‐gene phylogenies

inferred← citeshttps://doi.org/10.1111/eva.12269openalex-snapshot

Nonadaptive radiation in damselflies

inferred← citeshttps://doi.org/10.1111/evo.12436openalex-snapshot

ASYMMETRIC MISMATCH IN STRAIN-SPECIFIC GENITAL MORPHOLOGY CAUSES INCREASED HARM TODROSOPHILAFEMALES

inferred← citeshttps://doi.org/10.1111/evo.12463openalex-snapshot

A METHOD FOR ASSESSING PHYLOGENETIC LEAST SQUARES MODELS FOR SHAPE AND OTHER HIGH-DIMENSIONAL MULTIVARIATE DATA

inferred← citeshttps://doi.org/10.1111/evo.12596openalex-snapshot

Permutation tests for phylogenetic comparative analyses of high-dimensional shape data: What you shuffle matters

inferred← citeshttps://doi.org/10.1111/evo.12642openalex-snapshot

Relationships of diversity, disparity, and their evolutionary rates in squirrels (Sciuridae)

inferred← citeshttps://doi.org/10.1111/evo.12894openalex-snapshot

Tightly congruent bursts of lineage and phenotypic diversification identified in a continental ant radiation

inferred← citeshttps://doi.org/10.1111/evo.12939openalex-snapshot

Linking intra- and interspecific assortative mating: Consequences for asymmetric sexual isolation

inferred← citeshttps://doi.org/10.1111/evo.13315openalex-snapshot

Mechanical and tactile incompatibilities cause reproductive isolation between two young damselfly species

inferred← citeshttps://doi.org/10.1111/evo.13670openalex-snapshot

Adaptive radiation in labrid fishes: A central role for functional innovations during 65 My of relentless diversification

inferred← citeshttps://doi.org/10.1111/evo.13815openalex-snapshot

Head‐turning morphologies: Evolution of shape diversity in the mammalian atlas–axis complex

inferred← citeshttps://doi.org/10.1111/evo.13971openalex-snapshot

Decoupled jaws promote trophic diversity in cichlid fishes

inferred← citeshttps://doi.org/10.1111/j.1420-9101.2010.02000.xopenalex-snapshot

Divergent patterns of diversification in courtship and genitalic characters of Timema walking‐sticks

inferred← citeshttps://doi.org/10.1111/j.1420-9101.2011.02284.xopenalex-snapshot

Sexual selection and temporal phenotypic variation in a damselfly population

inferred← citeshttps://doi.org/10.1111/j.1420-9101.2011.02418.xopenalex-snapshot

Regional divergence and mosaic spatial distribution of two closely related damselfly species (Enallagma hageniandEnallagma ebrium)

inferred← citeshttps://doi.org/10.1111/j.1463-6409.2011.00500.xopenalex-snapshot

Evolution ofCryptocephalusleaf beetles related toC. sericeus(Coleoptera: Chrysomelidae) and the role of hybridization in generating species mtDNA paraphyly

inferred← citeshttps://doi.org/10.1111/j.1469-7998.2009.00620.xopenalex-snapshot

Geometric morphometrics: recent applications to the study of evolution and development

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2008.00527.xopenalex-snapshot

THE CORRELATED EVOLUTION OF THREE-DIMENSIONAL REPRODUCTIVE STRUCTURES BETWEEN MALE AND FEMALE DAMSELFLIES

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2008.00528.xopenalex-snapshot

STATIC ALLOMETRY AND ANIMAL GENITALIA

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2008.00557.xopenalex-snapshot

MODELING THREE-DIMENSIONAL MORPHOLOGICAL STRUCTURES USING SPHERICAL HARMONICS

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2009.00649.xopenalex-snapshot

A GENERAL FRAMEWORK FOR THE ANALYSIS OF PHENOTYPIC TRAJECTORIES IN EVOLUTIONARY STUDIES

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2010.01085.xopenalex-snapshot

A ROLE FOR LEARNING IN POPULATION DIVERGENCE OF MATE PREFERENCES

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2010.01138.xopenalex-snapshot

SPECIES RECOGNITION AND PATTERNS OF POPULATION VARIATION IN THE REPRODUCTIVE STRUCTURES OF A DAMSELFLY GENUS

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2011.01411.xopenalex-snapshot

SEXUAL SELECTION AND THE EVOLUTION OF GENITAL SHAPE AND COMPLEXITY IN WATER STRIDERS

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2011.01464.xopenalex-snapshot

SEXUAL CONFLICT AND ANTAGONISTIC COEVOLUTION ACROSS WATER STRIDER POPULATIONS

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2011.01469.xopenalex-snapshot

STRONG ASYMMETRY IN THE RELATIVE STRENGTHS OF PREZYGOTIC AND POSTZYGOTIC BARRIERS BETWEEN TWO DAMSELFLY SISTER SPECIES

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2011.01509.xopenalex-snapshot

EVIDENCE FOR STABILIZING SELECTION AND SLOW DIVERGENT EVOLUTION OF MALE GENITALIA IN A MILLIPEDE (ANTICHIROPUS VARIABILIS)

inferred← citeshttps://doi.org/10.1111/j.1558-5646.2011.01557.xopenalex-snapshot

RATES OF MORPHOLOGICAL EVOLUTION ARE CORRELATED WITH SPECIES RICHNESS IN SALAMANDERS

inferred← citeshttps://doi.org/10.1111/j.1749-6632.2009.04573.xopenalex-snapshot

Intralocus Sexual Conflict

inferred← citeshttps://doi.org/10.1111/jeb.12274openalex-snapshot

Genetic divergence predicts reproductive isolation in damselflies

inferred← citeshttps://doi.org/10.1111/jeb.12300openalex-snapshot

Renewed diversification is associated with new ecological opportunity in the N eotropical turtle ants

inferred← citeshttps://doi.org/10.1111/jeb.12955openalex-snapshot

The function and evolution of male and female genitalia in Phyllophaga Harris scarab beetles (Coleoptera: Scarabaeidae)

inferred← citeshttps://doi.org/10.1111/jeb.13302openalex-snapshot

Female mate preferences on high‐dimensional shape variation for male species recognition traits

inferred← citeshttps://doi.org/10.1111/jeb.13503openalex-snapshot

The evolution of female genitalia

inferred← citeshttps://doi.org/10.1111/jeb.13913openalex-snapshot

Evolution of male genitalia in the Drosophila repleta species group (Diptera: Drosophilidae)

inferred← citeshttps://doi.org/10.1111/jeb.13934openalex-snapshot

A holocenic and dynamic hybrid zone between two cactophilic Drosophila species in a coastal lowland plain of the Brazilian Atlantic Forest

inferred← citeshttps://doi.org/10.1111/zsc.12480openalex-snapshot

Congruence between molecular and morphological systematics of Alpine non‐biting midges (Chironomidae, Diamesinae)

inferred← citeshttps://doi.org/10.1134/s0013873811010027openalex-snapshot

The “lock-and-key” hypothesis as related to the ionizing radiation effect on genital cuticular structures in the large fruit-tree tortrix Archips podana (Lepidoptera, Tortricidae)

inferred← citeshttps://doi.org/10.1146/annurev-ecolsys-110218-024555openalex-snapshot

Phylogenetic Comparative Methods and the Evolution of Multivariate Phenotypes

inferred← citeshttps://doi.org/10.1146/annurev-ento-011613-161958openalex-snapshot

Freshwater Biodiversity and Aquatic Insect Diversification

inferred← citeshttps://doi.org/10.1163/18759866-08703001openalex-snapshot

Sexual behaviour and morphological variation in the millipede Megaphyllum bosniense (Verhoeff, 1897)

inferred← citeshttps://doi.org/10.11646/zootaxa.3815.2.1openalex-snapshot

The phallus in Tettigoniidae (Insecta: Orthoptera: Ensifera): revision of morphology and terminology, and discussion on its taxonomic importance and evolution

inferred← citeshttps://doi.org/10.1186/1471-2148-13-151openalex-snapshot

Hydrocarbon divergence and reproductive isolation in Timema stick insects

inferred← citeshttps://doi.org/10.1186/1471-2148-9-234openalex-snapshot

Inferring speciation modes in a clade of Iberian chafers from rates of morphological evolution in different character systems

inferred← citeshttps://doi.org/10.1186/s12862-014-0255-3openalex-snapshot

The role of aedeagus size and shape in failed mating interactions among recently diverged taxa in the Drosophila mojavensisspecies cluster

inferred← citeshttps://doi.org/10.1186/s12862-018-1305-zopenalex-snapshot

Alpha shapes: determining 3D shape complexity across morphologically diverse structures

inferred← citeshttps://doi.org/10.1186/s12915-021-01022-3openalex-snapshot

Sensory adaptations reshaped intrinsic factors underlying morphological diversification in bats

inferred← citeshttps://doi.org/10.1371/journal.pone.0013870openalex-snapshot

Biomechanical Consequences of Rapid Evolution in the Polar Bear Lineage

inferred← citeshttps://doi.org/10.1371/journal.pone.0026700openalex-snapshot

Contrasting Ecosystem-Effects of Morphologically Similar Copepods

inferred← citeshttps://doi.org/10.1371/journal.pone.0080531openalex-snapshot

Climate-Induced Range Shifts and Possible Hybridisation Consequences in Insects

inferred← citeshttps://doi.org/10.1371/journal.pone.0098536openalex-snapshot

The Evolution of Morphospace in Phytophagous Scarab Chafers: No Competition - No Divergence?

inferred← citeshttps://doi.org/10.1371/journal.pone.0128755openalex-snapshot

The Functional Significance of Chiral Genitalia: Patterns of Asymmetry, Functional Morphology and Mating Success in the Praying Mantis Ciulfina baldersoni

inferred← citeshttps://doi.org/10.1371/journal.pone.0137181openalex-snapshot

Biomechanical Diversity of Mating Structures among Harvestmen Species Is Consistent with a Spectrum of Precopulatory Strategies

inferred← citeshttps://doi.org/10.1371/journal.pone.0149673openalex-snapshot

Integrated Taxonomy and DNA Barcoding of Alpine Midges (Diptera: Chironomidae)

inferred← citeshttps://doi.org/10.15421/012426openalex-snapshot

Description of feather mite Proctophyllodes musicus (Acariformes, Proctophyllodidae) obtained from Turdus merula

inferred← citeshttps://doi.org/10.1890/11-1181.1openalex-snapshot

Climatic niche divergence or conservatism? Environmental niches and range limits in ecologically similar damselflies

inferred← citeshttps://doi.org/10.2139/ssrn.4822798openalex-snapshot

Regeneration of Realistic Grain Shapes Using Spherical Harmonic Analysis and Vector Quantized Time Series Generative Model

inferred← citeshttps://doi.org/10.3389/fevo.2021.642774openalex-snapshot

Computer Vision, Machine Learning, and the Promise of Phenomics in Ecology and Evolutionary Biology

inferred← citeshttps://doi.org/10.3917/flam.schil.2018.01.0281openalex-snapshot

Bibliographie

inferred← citeshttps://doi.org/10.4164/sptj.60.143openalex-snapshot

Particle Shape Generation using Spherical Harmonic Modeling and Principal Component Analysis and Application of the Discrete Element Method

inferred← citeshttps://doi.org/10.4164/sptj.62.579openalex-snapshot

A Method for Three-Dimensional Particle Shape Estimation and Generation

inferred← citeshttps://doi.org/10.4404/hystrix-24.1-6283openalex-snapshot

A field comes of age: geometric morphometrics in the 21 st century

inferred← citeshttps://doi.org/10.5325/jpennacadscie.93.1.0001openalex-snapshot

Environmental factors do not contribute significantly to shell shape in the Sprite Elimia, Elimia Proxima (Gastropoda: Pleuroceridae)

inferred← citeshttps://doi.org/10.5772/55195openalex-snapshot

Sexual Dimorphism Using Geometric Morphometric Approach

inferred← citeshttps://doi.org/10.6084/m9.figshare.14524705openalex-snapshot

Additional file 1 of Sensory adaptations reshaped intrinsic factors underlying morphological diversification in bats

inferred← citeshttps://doi.org/10.7554/elife.83426openalex-snapshot

Complex plumages spur rapid color diversification in kingfishers (Aves: Alcedinidae)

inferred← citeshttps://doi.org/10.7717/peerj.21266openalex-snapshot

Evolutionary divergence in a non-genitalic sexual contact character in the beetle genus Choleva

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