On the public citation graph
Inclusive fitness is an indispensable approximation for understanding organismal design
0 signed · 143 inferred · view as graph
Inferred edges come from two machine sources: sweeps of open repository metadata (mostly SSRN and Zenodo), and a bulk OpenAlex citation snapshot. Every row below is labelled with the one it came from. Neither is a complete citation index — absence here is not evidence of absence.
Is this your work? Sign in with ORCID and put your name on the
claims that are yours.
Reference health. See whether every reference in this work resolves to a
known work and none has been retracted — and get a dated CiteStamped
badge you can embed.
What this work says
Outgoing claims: the papers this work cites, builds on, or argues with.
inferredcites →https://doi.org/10.1002/ece3.6935openalex-snapshot
Extending the range of additivity in using inclusive fitness
inferredcites →https://doi.org/10.1006/anbe.1996.0020openalex-snapshot
The measurement and meaning of inclusive fitness
inferredcites →https://doi.org/10.1006/jtbi.1996.0075openalex-snapshot
How to Make a Kin Selection Model
inferredcites →https://doi.org/10.1007/s10539-013-9415-xopenalex-snapshot
Fitness, inclusive fitness, and optimization
inferredcites →https://doi.org/10.1007/s10539-016-9557-8openalex-snapshot
What’s wrong with evolutionary biology?
inferredcites →https://doi.org/10.1016/0003-3472(79)90028-9openalex-snapshot
The hawk-dove game played between relatives
inferredcites →https://doi.org/10.1016/0003-3472(79)90082-4openalex-snapshot
Behavioural ecology: An evolutionary approach
inferredcites →https://doi.org/10.1016/0022-5193(64)90038-4openalex-snapshot
The genetical evolution of social behaviour. I
inferredcites →https://doi.org/10.1016/0040-5809(78)90028-xopenalex-snapshot
Darwinian selection and “altruism”
inferredcites →https://doi.org/10.1016/j.cobeha.2016.09.015openalex-snapshot
What is inclusive fitness theory, and what is it for?
inferredcites →https://doi.org/10.1016/j.cobeha.2016.10.002openalex-snapshot
There is no inclusive fitness at the level of the individual
inferredcites →https://doi.org/10.1016/j.cub.2013.05.031openalex-snapshot
Adaptation and Inclusive Fitness
inferredcites →https://doi.org/10.1016/j.jtbi.2005.06.009openalex-snapshot
Optimization of inclusive fitness
inferredcites →https://doi.org/10.1016/j.jtbi.2007.03.015openalex-snapshot
The different limits of weak selection and the evolutionary dynamics of finite populations
inferredcites →https://doi.org/10.1016/j.jtbi.2015.04.031openalex-snapshot
Games among relatives revisited
inferredcites →https://doi.org/10.1016/j.jtbi.2015.06.039openalex-snapshot
Evolutionary dynamics of collective action in spatially structured populations
inferredcites →https://doi.org/10.1016/j.jtbi.2016.02.034openalex-snapshot
Hamilton׳s Rule in finite populations with synergistic interactions
inferredcites →https://doi.org/10.1038/2281218a0openalex-snapshot
Selfish and Spiteful Behaviour in an Evolutionary Model
inferredcites →https://doi.org/10.1038/298425a0openalex-snapshot
How not to measure inclusive fitness
inferredcites →https://doi.org/10.1038/318366a0openalex-snapshot
Kinship, reciprocity and synergism in the evolution of social behaviour
inferredcites →https://doi.org/10.1038/348027a0openalex-snapshot
Optimality theory in evolutionary biology
inferredcites →https://doi.org/10.1038/351619a0openalex-snapshot
Comparative methods for explaining adaptations
inferredcites →https://doi.org/10.1038/467661bopenalex-snapshot
Inclusive fitness is just bookkeeping
inferredcites →https://doi.org/10.1038/nature09335openalex-snapshot
Promiscuity and the evolutionary transition to complex societies
inferredcites →https://doi.org/10.1038/nature09831openalex-snapshot
Inclusive fitness theory and eusociality
inferredcites →https://doi.org/10.1038/ncomms15973openalex-snapshot
The evolution of host-symbiont dependence
inferredcites →https://doi.org/10.1038/nrg2903openalex-snapshot
The sociobiology of molecular systems
inferredcites →https://doi.org/10.1038/s41559-016-0057openalex-snapshot
Cooperation facilitates the colonization of harsh environments
inferredcites →https://doi.org/10.1038/s41559-017-0354-9openalex-snapshot
The meaning of intragenomic conflict
inferredcites →https://doi.org/10.1073/pnas.1100298108openalex-snapshot
Expanded social fitness and Hamilton's rule for kin, kith, and kind
inferredcites →https://doi.org/10.1073/pnas.1317588110openalex-snapshot
Limitations of inclusive fitness
inferredcites →https://doi.org/10.1073/pnas.1701805114openalex-snapshot
The general form of Hamilton’s rule makes no predictions and cannot be tested empirically
inferredcites →https://doi.org/10.1073/pnas.74.10.4542openalex-snapshot
How does selection reconcile individual advantage with the good of the group?
inferredcites →https://doi.org/10.1086/284016openalex-snapshot
Population Genetic Theory of Kin Selection. II. The Multiplicative Model
inferredcites →https://doi.org/10.1093/biosci/biv129openalex-snapshot
Inclusive Fitness Theory Becomes an End in Itself
inferredcites →https://doi.org/10.1093/bjps/axt016openalex-snapshot
Hamilton’s Rule and Its Discontents
inferredcites →https://doi.org/10.1093/oso/9780198546412.001.0001openalex-snapshot
The Comparative Method in Evolutionary Biology
inferredcites →https://doi.org/10.1093/oso/9780198733058.001.0001openalex-snapshot
The Philosophy of Social Evolution
inferredcites →https://doi.org/10.1093/qmath/12.1.78openalex-snapshot
ON AN INEQUALITY IN PARTIAL AVERAGES
inferredcites →https://doi.org/10.1098/rsos.160037openalex-snapshot
The causal meaning of Hamilton’s rule
inferredcites →https://doi.org/10.1098/rsos.170335openalex-snapshot
The inclusive fitness controversy: finding a way forward
inferredcites →https://doi.org/10.1098/rspb.1983.0077openalex-snapshot
The eleventh R. A. Fisher Memorial Lecture - Kin selection and altruism
inferredcites →https://doi.org/10.1098/rspb.2011.1465openalex-snapshot
The validity and value of inclusive fitness theory
inferredcites →https://doi.org/10.1098/rstb.2015.0085openalex-snapshot
There is no fitness but fitness, and the lineage is its bearer
inferredcites →https://doi.org/10.1111/evo.12701openalex-snapshot
Does evolution lead to maximizing behavior?
inferredcites →https://doi.org/10.1111/evo.12791openalex-snapshot
Regression, least squares, and the general version of inclusive fitness
inferredcites →https://doi.org/10.1111/evo.12980openalex-snapshot
Invasion fitness, inclusive fitness, and reproductive numbers in heterogeneous populations
inferredcites →https://doi.org/10.1111/evo.13160openalex-snapshot
Inclusive fitness in finite populations-effects of heterogeneity and synergy
inferredcites →https://doi.org/10.1111/j.1420-9101.2006.01258.xopenalex-snapshot
Social semantics: altruism, cooperation, mutualism, strong reciprocity and group selection
inferredcites →https://doi.org/10.1111/j.1420-9101.2007.01458.xopenalex-snapshot
Social semantics: how useful has group selection been?
inferredcites →https://doi.org/10.1111/j.1420-9101.2008.01681.xopenalex-snapshot
Capturing the superorganism: a formal theory of group adaptation
inferredcites →https://doi.org/10.1111/j.1420-9101.2011.02236.xopenalex-snapshot
The genetical theory of kin selection
inferredcites →https://doi.org/10.1111/j.1558-5646.1992.tb02045.xopenalex-snapshot
A GENERAL MODEL FOR KIN SELECTION
inferredcites →https://doi.org/10.1111/j.1558-5646.2009.00929.xopenalex-snapshot
THE SOCIAL ORGANISM: CONGRESSES, PARTIES, AND COMMITTEES
inferredcites →https://doi.org/10.1111/jeb.12808openalex-snapshot
Hamilton's rule, inclusive fitness maximization, and the goal of individual behaviour in symmetric two‐player games
inferredcites →https://doi.org/10.1126/science.1156108openalex-snapshot
Ancestral Monogamy Shows Kin Selection Is Key to the Evolution of Eusociality
inferredcites →https://doi.org/10.1146/annurev.es.09.110178.002313openalex-snapshot
Group Selection, Altruism, and the Levels of Organization of Life
inferredcites →https://doi.org/10.1371/journal.pbio.1002134openalex-snapshot
Inclusive Fitness Theorizing Invokes Phenomena That Are Not Relevant for the Evolution of Eusociality
inferredcites →https://doi.org/10.1515/9780691206820openalex-snapshot
Foundations of Social Evolution
inferredcites →https://doi.org/10.1515/9781400847242openalex-snapshot
Genetic Structure and Selection in Subdivided Populations (MPB-40)
inferredcites →https://doi.org/10.1515/9781400866564openalex-snapshot
Social Evolution and Inclusive Fitness Theory
inferredcites →https://doi.org/10.1890/0012-9658(2006)87[1445:soefdd]2.0.co;2openalex-snapshot
STRENGTH OF EVIDENCE FOR DENSITY DEPENDENCE IN ABUNDANCE TIME SERIES OF 1198 SPECIES
inferredcites →https://doi.org/10.5962/bhl.title.2092openalex-snapshot
The descent of man, and selection in relation to sex
What points at this work
Incoming claims: where this work appears in other papers’ reference lists and arguments.
inferred← citeshttps://doi.org/10.2139/ssrn.4815379openalex-extracted
Syntax Before Words—The Function-First Approach to the Evolution of Proto-Language
inferred← citeshttps://doi.org/10.1002/ece3.6935openalex-snapshot
Extending the range of additivity in using inclusive fitness
inferred← citeshttps://doi.org/10.1007/978-3-031-22028-9_35openalex-snapshot
Genes and Organisms in the Legacy of the Modern Synthesis
inferred← citeshttps://doi.org/10.1007/978-3-031-22028-9_5openalex-snapshot
The Creativity of Natural Selection and the Creativity of Organisms: Their Roles in Traditional Evolutionary Theory and Some Proposed Extensions
inferred← citeshttps://doi.org/10.1007/s10539-025-10001-9openalex-snapshot
Internal conflicts and the measurement of evolutionary individuality
inferred← citeshttps://doi.org/10.1007/s13194-021-00363-8openalex-snapshot
Misconceptions, conceptual pluralism, and conceptual toolkits: bringing the philosophy of science to the teaching of evolution
inferred← citeshttps://doi.org/10.1007/s13235-025-00638-2openalex-snapshot
Beyond the Classical Hamilton’s Rule: State Distribution Asymmetry and the Dynamics of Altruism
inferred← citeshttps://doi.org/10.1007/s13752-021-00381-yopenalex-snapshot
The Scapegoat Mechanism in Human Evolution: An Analysis of René Girard’s Hypothesis on the Process of Hominization
inferred← citeshttps://doi.org/10.1016/j.shpsa.2021.11.008openalex-snapshot
Hamilton's rule: A non-causal explanation?
inferred← citeshttps://doi.org/10.1016/j.shpsa.2026.102205openalex-snapshot
On the general forms of Hamilton’s rule derived from regression models of social evolution
inferred← citeshttps://doi.org/10.1016/j.tpb.2024.02.008openalex-snapshot
Evolution of spite versus evolution of altruism through a disbandment mechanism
inferred← citeshttps://doi.org/10.1017/9781009019644openalex-snapshot
Inclusive Fitness and Kin Selection
inferred← citeshttps://doi.org/10.1038/s41467-022-31545-4openalex-snapshot
Multiple social encounters can eliminate Crozier’s paradox and stabilise genetic kin recognition
inferred← citeshttps://doi.org/10.1038/s42003-023-05260-9openalex-snapshot
Inclusive fitness forces of selection in an age-structured population
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0001openalex-snapshot
Making room for maladaptation
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0002openalex-snapshot
Natural selection through genetics
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0003openalex-snapshot
Population ecology of natural selection
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0004openalex-snapshot
Maladaptation in social behaviour
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0005openalex-snapshot
Maladaptation within the body
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0006openalex-snapshot
Maladaptive transitions in complexity
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0007openalex-snapshot
Deep origins of maladaptation
inferred← citeshttps://doi.org/10.1093/9780191966767.003.0008openalex-snapshot
Revisiting the design argument
inferred← citeshttps://doi.org/10.1093/9780191966767.005.0001openalex-snapshot
Appendix A simple mathematical model of maladaptation
inferred← citeshttps://doi.org/10.1093/icb/icaa029openalex-snapshot
The Centrality of Ancestral Grandmothering in Human Evolution
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.001.0001openalex-snapshot
Blue-Footed Boobies
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.002.0009openalex-snapshot
List of Common and Systematic Names
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.002.0010openalex-snapshot
Advanced Praise for Blue-Footed Boobies
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0002openalex-snapshot
Two Approaches to Controlling and Killing Siblings
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0003openalex-snapshot
Beating Siblings into Submission
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0004openalex-snapshot
From Wild Violence to Courtly Rituals
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0005openalex-snapshot
To Kill or Not to Kill
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0006openalex-snapshot
Are Parents Okay with Sibling Bullying?
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0007openalex-snapshot
Bullying and Lifelong Scars
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0008openalex-snapshot
Happy Marriages with Blue Feet
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0009openalex-snapshot
Cheating, Infanticide, and Egg-Dumping
inferred← citeshttps://doi.org/10.1093/oso/9780197629840.003.0010openalex-snapshot
Are Humans Similar?
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.001.0001openalex-snapshot
Domains and Major Transitions of Social Evolution
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0001openalex-snapshot
A punctuated history of understanding social adaptation
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0002openalex-snapshot
The gene’s eye view that forged a neo-Darwinian synthesis
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0003openalex-snapshot
A reappraisal of progress in evolution
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0004openalex-snapshot
Necessary and sufficient conditions for major evolutionary transitions
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0005openalex-snapshot
Inclusive fitness as driver of cooperation for mutual benefit
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0006openalex-snapshot
The multicellular organisms and colonial superorganisms
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0007openalex-snapshot
The free-living prokaryotic and eukaryotic cells
inferred← citeshttps://doi.org/10.1093/oso/9780198746171.003.0008openalex-snapshot
Adaptation, control information, and the human condition
inferred← citeshttps://doi.org/10.1093/pnasnexus/pgae131openalex-snapshot
Nonlinear social evolution and the emergence of collective action
inferred← citeshttps://doi.org/10.1098/rspb.2019.1060openalex-snapshot
What life is for: a commentary on Fromhage and Jennions
inferred← citeshttps://doi.org/10.1098/rspb.2023.1310openalex-snapshot
How to make an inclusive-fitness model
inferred← citeshttps://doi.org/10.1101/2023.10.02.560418openalex-snapshot
Cochlear tuning characteristics arise from temporal prediction of natural sounds
inferred← citeshttps://doi.org/10.1101/cshperspect.a041667openalex-snapshot
Calculating Relatedness: A Pedigree of Definitions
inferred← citeshttps://doi.org/10.1111/1365-2656.13839openalex-snapshot
Social network analysis reveals context‐dependent kin relationships in wild sulphur‐crested cockatoos Cacatua galerita
inferred← citeshttps://doi.org/10.1134/s0006297922120094openalex-snapshot
Tracing Slow Phenoptosis to the Prenatal Stage in Social Vertebrates
inferred← citeshttps://doi.org/10.1177/17456916231201512openalex-snapshot
The Effect of Income and Wealth on Behavioral Strategies, Personality Traits, and Preferences
inferred← citeshttps://doi.org/10.1371/journal.pcbi.1010612openalex-snapshot
Emergent multilevel selection in a simple spatial model of the evolution of altruism
inferred← citeshttps://doi.org/10.31234/osf.io/2r3efopenalex-snapshot
Optimal resource allocation and its consequences on behavioral strategies, personality traits and preferences
inferred← citeshttps://doi.org/10.3389/fevo.2023.1229093openalex-snapshot
The semantics of stability: evolutionarily stable strategy in biology and economics literature
inferred← citeshttps://doi.org/10.3389/fpsyt.2023.1110420openalex-snapshot
The evolution of personality disorders: A review of proposals
inferred← citeshttps://doi.org/10.3390/encyclopedia4010029openalex-snapshot
The Evolution of Human Social Behavior
inferred← citeshttps://doi.org/10.70675/262719dbz1132z4de5z9e4bzae9ea9ab6b0copenalex-snapshot
A mechanistic view of the evolution of cell diversity : or when constraints meet chance and necessity in life’s struggle for multiplication
Editorial review for independent work. ICSAC — the Institute for Complexity
Science and Advanced Computing — runs a structured editorial panel for researchers
publishing outside journals, and publishes on CiteStamp rails. Submission is free
and goes through ORCID: icsacinstitute.org/submit.