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Solving the missing heritability problem
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What this work says
Outgoing claims: the papers this work cites, builds on, or argues with.
inferredcites →https://doi.org/10.1016/j.ajhg.2010.11.011openalex-snapshot
GCTA: A Tool for Genome-wide Complex Trait Analysis
inferredcites →https://doi.org/10.1016/j.ajhg.2013.10.005openalex-snapshot
Inference of the Genetic Architecture Underlying BMI and Height with the Use of 20,240 Sibling Pairs
inferredcites →https://doi.org/10.1016/j.ajhg.2019.06.006openalex-snapshot
Comparing Within- and Between-Family Polygenic Score Prediction
inferredcites →https://doi.org/10.1016/j.ssresearch.2013.10.004openalex-snapshot
What can we learn from twin studies? A comprehensive evaluation of the equal environments assumption
inferredcites →https://doi.org/10.1038/nature07331openalex-snapshot
Genes mirror geography within Europe
inferredcites →https://doi.org/10.1038/nature08494openalex-snapshot
Finding the missing heritability of complex diseases
inferredcites →https://doi.org/10.1038/ng.1074openalex-snapshot
Differential confounding of rare and common variants in spatially structured populations
inferredcites →https://doi.org/10.1038/ng.2620openalex-snapshot
FaST-LMM-Select for addressing confounding from spatial structure and rare variants
inferredcites →https://doi.org/10.1038/ng.2876openalex-snapshot
Advantages and pitfalls in the application of mixed-model association methods
inferredcites →https://doi.org/10.1038/ng.3390openalex-snapshot
Genetic variance estimation with imputed variants finds negligible missing heritability for human height and body mass index
inferredcites →https://doi.org/10.1038/ng.3865openalex-snapshot
Reevaluation of SNP heritability in complex human traits
inferredcites →https://doi.org/10.1038/ng.3941openalex-snapshot
Concepts, estimation and interpretation of SNP-based heritability
inferredcites →https://doi.org/10.1038/ng.548openalex-snapshot
Variance component model to account for sample structure in genome-wide association studies
inferredcites →https://doi.org/10.1038/ng.608openalex-snapshot
Common SNPs explain a large proportion of the heritability for human height
inferredcites →https://doi.org/10.1038/nrg2809openalex-snapshot
Missing heritability and strategies for finding the underlying causes of complex disease
inferredcites →https://doi.org/10.1038/s41467-017-02088-wopenalex-snapshot
Structural absorption by barbule microstructures of super black bird of paradise feathers
inferredcites →https://doi.org/10.1038/s41467-018-06964-xopenalex-snapshot
A homozygous loss-of-function mutation leading to CYBC1 deficiency causes chronic granulomatous disease
inferredcites →https://doi.org/10.1038/s41588-018-0147-3openalex-snapshot
Gene discovery and polygenic prediction from a genome-wide association study of educational attainment in 1.1 million individuals
inferredcites →https://doi.org/10.1038/s41588-018-0178-9openalex-snapshot
Relatedness disequilibrium regression estimates heritability without environmental bias
inferredcites →https://doi.org/10.1038/s41588-019-0465-0openalex-snapshot
Accurate estimation of SNP-heritability from biobank-scale data irrespective of genetic architecture
inferredcites →https://doi.org/10.1038/s41588-021-00997-7openalex-snapshot
Assessing the contribution of rare variants to complex trait heritability from whole-genome sequence data
inferredcites →https://doi.org/10.1073/pnas.1119675109openalex-snapshot
The mystery of missing heritability: Genetic interactions create phantom heritability
inferredcites →https://doi.org/10.1101/048181openalex-snapshot
Subtle stratification confounds estimates of heritability from rare variants
inferredcites →https://doi.org/10.1126/science.aan6877openalex-snapshot
The nature of nurture: Effects of parental genotypes
inferredcites →https://doi.org/10.1371/journal.pbio.2002985openalex-snapshot
A population genetic interpretation of GWAS findings for human quantitative traits
inferredcites →https://doi.org/10.1371/journal.pgen.0020041openalex-snapshot
Assumption-Free Estimation of Heritability from Genome-Wide Identity-by-Descent Sharing between Full Siblings
inferredcites →https://doi.org/10.1371/journal.pgen.0020190openalex-snapshot
Population Structure and Eigenanalysis
inferredcites →https://doi.org/10.1371/journal.pgen.1004269openalex-snapshot
Statistical Power to Detect Genetic (Co)Variance of Complex Traits Using SNP Data in Unrelated Samples
inferredcites →https://doi.org/10.1375/136905202320906219openalex-snapshot
The Swedish Twin Registry in the Third Millennium
inferredcites →https://doi.org/10.1375/136905202762342026openalex-snapshot
Variance Components Models for Gene–Environment Interaction in Twin Analysis
inferredcites →https://doi.org/10.1375/136905203770326402openalex-snapshot
Heritability of Adult Body Height: A Comparative Study of Twin Cohorts in Eight Countries
inferredcites →https://doi.org/10.1530/ey.16.14.15openalex-snapshot
Recovery of trait heritability from whole genome sequence data
inferredcites →https://doi.org/10.1534/genetics.119.301859openalex-snapshot
Complex Trait Prediction from Genome Data: Contrasting EBV in Livestock to PRS in Humans
inferredcites →https://doi.org/10.2307/2529912openalex-snapshot
Introduction to Quantitative Genetics.
inferredcites →https://doi.org/10.7554/elife.39702openalex-snapshot
Polygenic adaptation on height is overestimated due to uncorrected stratification in genome-wide association studies
inferredcites →https://doi.org/10.7554/elife.39725openalex-snapshot
Reduced signal for polygenic adaptation of height in UK Biobank
inferredcites →https://doi.org/10.7554/elife.48376openalex-snapshot
Variable prediction accuracy of polygenic scores within an ancestry group
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Incoming claims: where this work appears in other papers’ reference lists and arguments.
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Genetic Liability for Schizophrenia and Childhood Psychopathology in the General Population
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Heritability of musculoskeletal pain and pain sensitivity phenotypes: 2 generations of the Raine Study
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Evidence of positive and negative selection associated with DNA methylation
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Genetic influences on neurodevelopmental disorders and their overlap with co-occurring conditions in childhood and adolescence: A meta-analysis
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DrFARM: Identification and inference for pleiotropic gene in GWAS
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Estimation of indirect genetic effects and heritability under assortative mating
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Genetic liability to sedentary behavior increases the risk of cardiovascular disease incidence: Evidence from the FinnGen cohort with 293,250 individuals
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Explicitly modeling genetic ancestry to improve polygenic prediction accuracy for height in a large, admixed cohort of US Latinos: Findings from HCHS/SOL
inferred← citeshttps://doi.org/10.1101/2025.04.11.648325openalex-snapshot
Biologically-informed Interpretable Deep Learning Framework for Phenotype Prediction and Gene Interaction Detection
inferred← citeshttps://doi.org/10.1101/2025.04.20.649385openalex-snapshot
Heritability of human lifespan is about 50% when confounding factors are addressed
inferred← citeshttps://doi.org/10.1101/2025.10.08.680992openalex-snapshot
Beyond years of schooling: Shifting genetic influences across educational milestones in two Norwegian cohorts
inferred← citeshttps://doi.org/10.1109/tit.2025.3529457openalex-snapshot
Optimal Estimation of the Null Distribution in Large-Scale Inference
inferred← citeshttps://doi.org/10.1111/eva.13264openalex-snapshot
Meta‐analysis: Congruence of genomic and phenotypic differentiation across diverse natural study systems
inferred← citeshttps://doi.org/10.1111/jcmm.16433openalex-snapshot
Aryl hydrocarbon receptor (Ahr)‐dependent Il‐22 expression by type 3 innate lymphoid cells control of acute joint inflammation
inferred← citeshttps://doi.org/10.1111/jcpp.13654openalex-snapshot
On the importance of parenting in externalizing disorders: an evaluation of indirect genetic effects in families
inferred← citeshttps://doi.org/10.1111/jcpp.14099openalex-snapshot
Research Review: A review of the past decade of family and genomic studies on adolescent mental health
inferred← citeshttps://doi.org/10.1111/mec.16367openalex-snapshot
Polygenic adaptation and negative selection across traits, years and environments in a long‐lived plant species (Pinus pinaster Ait., Pinaceae)
inferred← citeshttps://doi.org/10.1136/bjsports-2024-109491openalex-snapshot
Genetic liability to sedentary behaviour and cardiovascular disease incidence in the FinnGen and HUNT cohorts
inferred← citeshttps://doi.org/10.1146/annurev-biodatasci-103123-095355openalex-snapshot
Genetic Studies Through the Lens of Gene Networks
inferred← citeshttps://doi.org/10.1146/annurev-clinpsy-081219-091234openalex-snapshot
What Do We Know About the Genetic Architecture of Psychopathology?
inferred← citeshttps://doi.org/10.1146/annurev-genet-021920-102037openalex-snapshot
What Has a Century of Quantitative Genetics Taught Us About Nature's Genetic Tool Kit?
inferred← citeshttps://doi.org/10.1146/annurev-genom-120219-074937openalex-snapshot
The Genetics of Epilepsy
inferred← citeshttps://doi.org/10.1146/annurev-psych-052220-103822openalex-snapshot
“Reports of My Death Were Greatly Exaggerated”: Behavior Genetics in the Postgenomic Era
inferred← citeshttps://doi.org/10.1155/2022/7843990openalex-snapshot
Maximal Information Coefficient-Based Testing to Identify Epistasis in Case-Control Association Studies
inferred← citeshttps://doi.org/10.1177/0004867420979353openalex-snapshot
The 2020 Royal Australian and New Zealand College of Psychiatrists clinical practice guidelines for mood disorders
inferred← citeshttps://doi.org/10.1177/17456916211041602openalex-snapshot
The “Golden Age” of Behavior Genetics?
inferred← citeshttps://doi.org/10.1177/23312165251320156openalex-snapshot
Genetic and Environmental Contributions to Age-Related Hearing Loss: Results from a Longitudinal Twin Study
inferred← citeshttps://doi.org/10.1186/s12284-023-00621-8openalex-snapshot
Identification of Novel Quantitative Trait Loci for Culm Thickness of Rice Derived from Strong-Culm Landrace in Japan, Omachi
inferred← citeshttps://doi.org/10.1186/s12711-022-00763-1openalex-snapshot
Failing the four-gamete test enables exact phasing: the Corners’ Algorithm
inferred← citeshttps://doi.org/10.1186/s12711-025-00993-zopenalex-snapshot
Single-variant genome-wide association study and regional heritability mapping of protein efficiency and performance traits in Large White pigs
inferred← citeshttps://doi.org/10.1186/s12859-022-04987-2openalex-snapshot
CLIN_SKAT: an R package to conduct association analysis using functionally relevant variants
inferred← citeshttps://doi.org/10.1186/s12859-023-05451-5openalex-snapshot
A knowledge graph approach to predict and interpret disease-causing gene interactions
inferred← citeshttps://doi.org/10.1186/s12863-023-01155-0openalex-snapshot
Gene-based burden scores identify rare variant associations for 28 blood biomarkers
inferred← citeshttps://doi.org/10.1186/s12967-021-03090-zopenalex-snapshot
Integrative eQTL-weighted hierarchical Cox models for SNP-set based time-to-event association studies
inferred← citeshttps://doi.org/10.1186/s13059-021-02392-1openalex-snapshot
Leveraging breeding programs and genomic data in Norway spruce (Picea abies L. Karst) for GWAS analysis
inferred← citeshttps://doi.org/10.1186/s13059-022-02697-9openalex-snapshot
Open problems in human trait genetics
inferred← citeshttps://doi.org/10.1186/s13059-026-04140-9openalex-snapshot
UK Biobank whole-genome sequencing reveals robust contributions of rare variants to complex-trait heritability
inferred← citeshttps://doi.org/10.1186/s13062-024-00574-yopenalex-snapshot
Integrating multi-layered biological priors to improve genomic prediction accuracy in beef cattle
inferred← citeshttps://doi.org/10.1186/s13064-026-00236-xopenalex-snapshot
Modifier gene identification reveals therapeutic pathways for adult and childhood dementias
inferred← citeshttps://doi.org/10.1249/mss.0000000000002981openalex-snapshot
A Polygenic Risk Score for Hand Grip Strength Predicts Muscle Strength and Proximal and Distal Functional Outcomes among Older Women
inferred← citeshttps://doi.org/10.1249/mss.0000000000004019openalex-snapshot
Evaluating a Genome-Wide Polygenic Score for Handgrip Strength and Its Interplay with Leisure-Time Physical Activity Across the IGEMS Twin Cohorts
inferred← citeshttps://doi.org/10.1371/journal.pbio.3002847openalex-snapshot
Unifying approaches from statistical genetics and phylogenetics for mapping phenotypes in structured populations
inferred← citeshttps://doi.org/10.1371/journal.pgen.1008791openalex-snapshot
The genetic architecture of the maize progenitor, teosinte, and how it was altered during maize domestication
inferred← citeshttps://doi.org/10.1371/journal.pgen.1009762openalex-snapshot
Average semivariance yields accurate estimates of the fraction of marker-associated genetic variance and heritability in complex trait analyses
inferred← citeshttps://doi.org/10.1371/journal.pgen.1010567openalex-snapshot
Using epigenomics to understand cellular responses to environmental influences in diseases
inferred← citeshttps://doi.org/10.1371/journal.pgen.1010638openalex-snapshot
The identification of mediating effects using genome-based restricted maximum likelihood estimation
inferred← citeshttps://doi.org/10.1371/journal.pgen.1010863openalex-snapshot
Artificial selection reveals complex genetic architecture of shoot branching and its response to nitrate supply in Arabidopsis
inferred← citeshttps://doi.org/10.1371/journal.pgen.1011730openalex-snapshot
A mathematical framework for the quantitative analysis of genetic buffering
inferred← citeshttps://doi.org/10.1371/journal.pgen.1012310openalex-snapshot
Beyond years of schooling: Genetic associations across educational milestones in two Norwegian cohorts
inferred← citeshttps://doi.org/10.1371/journal.pone.0263390openalex-snapshot
Evaluating the detection ability of a range of epistasis detection methods on simulated data for pure and impure epistatic models
inferred← citeshttps://doi.org/10.1371/journal.pone.0294896openalex-snapshot
Cohort profile: Genetic data in the German Socio-Economic Panel Innovation Sample (SOEP-G)
inferred← citeshttps://doi.org/10.1371/journal.pone.0314502openalex-snapshot
TRIO RVEMVS: A Bayesian framework for rare variant association analysis with expectation-maximization variable selection using family trio data
inferred← citeshttps://doi.org/10.1371/journal.pone.0348023openalex-snapshot
Quantitative genetic analysis of respiratory function and related traits in Bulldogs, French Bulldogs and Pugs
inferred← citeshttps://doi.org/10.1515/sagmb-2022-0034openalex-snapshot
Bayesian LASSO for population stratification correction in rare haplotype association studies
inferred← citeshttps://doi.org/10.1530/eor-22-0026openalex-snapshot
Idiopathic scoliosis: a systematic review and meta-analysis of heritability
inferred← citeshttps://doi.org/10.15302/j-qb-021-0249openalex-snapshot
Advances and challenges in quantitative delineation of the genetic architecture of complex traits
inferred← citeshttps://doi.org/10.17513/mjpfi.13501openalex-snapshot
GENETIC RISK FACTOR FOR IDIOPATHIC SCOLIOSIS
inferred← citeshttps://doi.org/10.18261/issn.1504-291x-2020-01-05openalex-snapshot
Fra sosiobiologi til biososiologi
inferred← citeshttps://doi.org/10.18632/aging.203262openalex-snapshot
Sex differences in genetic and environmental influences on frailty and its relation to body mass index and education
inferred← citeshttps://doi.org/10.18865/ed.34.2.103openalex-snapshot
Intracranial Volume Is Driven by Both Genetics and Early Life Exposures: The SOL-INCA-MRI Study
inferred← citeshttps://doi.org/10.20377/jfr-381openalex-snapshot
Trait-specific testing of the equal environment assumption: The case of school grades and upper secondary school attendance
inferred← citeshttps://doi.org/10.2147/opto.s226595openalex-snapshot
Monitoring and Management of the Patient with Stargardt Disease
inferred← citeshttps://doi.org/10.26508/lsa.202000902openalex-snapshot
A novel canis lupus familiaris reference genome improves variant resolution for use in breed-specific GWAS
inferred← citeshttps://doi.org/10.29220/csam.2022.29.1.065openalex-snapshot
Grid-based Gaussian process models for longitudinal genetic data
inferred← citeshttps://doi.org/10.29220/csam.2022.29.1.745openalex-snapshot
Grid-based Gaussian process models for longitudinal genetic data
inferred← citeshttps://doi.org/10.31235/osf.io/5a7vxopenalex-snapshot
The Genetic and Environmental Composition of Socioeconomic Status: A population-wide multi-method study in >170,000 Norwegian individuals
inferred← citeshttps://doi.org/10.31274/etd-20210609-139openalex-snapshot
Genotypes to Phenotypes: Genomic attributes that impact Genome-wide Associations in peanut (Arachis hypogaea. L).
inferred← citeshttps://doi.org/10.3233/jpd-212818openalex-snapshot
Insights on Genetic and Environmental Factors in Parkinson’s Disease from a Regional Swedish Case-Control Cohort
inferred← citeshttps://doi.org/10.3389/fbinf.2026.1800403openalex-snapshot
Polygenic modeling of genetic effects on both phenotypic mean and variance: distributional regression for BMI, blood and urine biomarkers in the UK Biobank
inferred← citeshttps://doi.org/10.3389/fcell.2021.801113openalex-snapshot
Gene-Based Testing of Interactions Using XGBoost in Genome-Wide Association Studies
inferred← citeshttps://doi.org/10.3389/fgene.2020.592461openalex-snapshot
Improved Detection of Potentially Pleiotropic Genes in Coronary Artery Disease and Chronic Kidney Disease Using GWAS Summary Statistics
inferred← citeshttps://doi.org/10.3389/fgene.2021.628192openalex-snapshot
KLF4, a Key Regulator of a Transitive Triplet, Acts on the TGF-β Signaling Pathway and Contributes to High-Altitude Adaptation of Tibetan Pigs
inferred← citeshttps://doi.org/10.3389/fgene.2021.709555openalex-snapshot
An Integrative Co-localization (INCO) Analysis for SNV and CNV Genomic Features With an Application to Taiwan Biobank Data
inferred← citeshttps://doi.org/10.3389/fgene.2021.801261openalex-snapshot
Testing Gene-Gene Interactions Based on a Neighborhood Perspective in Genome-wide Association Studies
inferred← citeshttps://doi.org/10.3389/fimmu.2022.975115openalex-snapshot
New insights on the role of human leukocyte antigen complex in primary biliary cholangitis
inferred← citeshttps://doi.org/10.3389/fncel.2023.1112405openalex-snapshot
Large-scale analyses of CAV1 and CAV2 suggest their expression is higher in post-mortem ALS brain tissue and affects survival
inferred← citeshttps://doi.org/10.3389/fpsyt.2026.1887517openalex-snapshot
Beyond the brain circuit: a biopsychosocial integrative review of addiction neuroscience, epigenetic mechanisms, and multidisciplinary treatment
inferred← citeshttps://doi.org/10.3389/fvets.2022.1045589openalex-snapshot
Genome-wide association study reveals novel candidate genes for litter size in Markhoz goats
inferred← citeshttps://doi.org/10.3390/genes11040464openalex-snapshot
Identification of Age-Specific and Common Key Regulatory Mechanisms Governing Eggshell Strength in Chicken Using Random Forests
inferred← citeshttps://doi.org/10.3390/genes11080892openalex-snapshot
Combining Random Forests and a Signal Detection Method Leads to the Robust Detection of Genotype-Phenotype Associations
inferred← citeshttps://doi.org/10.3390/genes13040646openalex-snapshot
Major Depressive Disorder: Existing Hypotheses about Pathophysiological Mechanisms and New Genetic Findings
inferred← citeshttps://doi.org/10.3390/ijms25021347openalex-snapshot
SumVg: Total Heritability Explained by All Variants in Genome-Wide Association Studies Based on Summary Statistics with Standard Error Estimates
inferred← citeshttps://doi.org/10.3390/ijms25031918openalex-snapshot
Advancements and Prospects of Genome-Wide Association Studies (GWAS) in Maize
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